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10.1016/s0140-6736(77)90612-2 Lancet

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We therefore propose that acidification of mutant supernatant (via secretion of acids and impaired urease activity) and changes to extracellular antioxidant composition circumvented the oxidation of L-DOPA, thereby preventing its use in melanin formation by gsh2 mutants

Journal of Applied Physiology, 110(3), 774-780